> Can it always tune up from below the note?
Read that whole site. This is an Engineering Project, NOT a useful tool. The students solved enough engineering problems to get a grade, but it isn't useful yet. It has poor range, poor resolution, poor noise rejection, chokes on pluck-noise, and moving the motor from one peg to the next is utterly impractical. Fine details like "always tune up" are not handled.
Anyway.....
Are we musicians or technicians?
Actually, I am a technician with no musical chops. But I can tune a dead-beat by ear. And if you have a pitch reference (fork or dialtone) that's all you need to tune a simple fretted instrument like a guitar. And making two notes harmonious is the basic skill of melody/harmony musicians. And one of the few things that separates people from "dumb" animals.